Jon Karlen is a partner in the technology group at Atlas Venture and focuses on investments in software and online services. Jon joined Atlas in early 2014 and sits on the boards of CustomMade, Fashion Project, Kinvey, PillPack, and Robin. Prior to joining Atlas, Jon was a partner at Flybridge Capital, where he led seed investments in disruptive software businesses including Open English and Digital Lumens. He also pioneered investments in Virtual Computer (acquired by Citrix), ZING Systems (acquired by Dell), and Restorando. Jon began his career at Greylock where as an EIR he developed an investment opportunity in Oat Systems, an RFID software company. Jon is a frequent contributor and advisor at local incubators, and founded the University Research and Entrepreneurship Symposium, a one day conference that connects investors with technology startups spun out of university research. He received an MBA from Stanford University and an AB from Harvard College.